Adjustment of breeding values for past and future inbreeding Paul VanRaden*, Lori Smith Animal Improvement Programs Laboratory Agricultural Research Service, USDA Beltsville, Maryland Selection and Mating Selection Rank animals from best to worst Decide which animals reproduce Mating Predict the merit of pairs Decide which animals are paired Select/Mate
Do both steps at once Better in theory but not practical ADSA 1999 (2) P. M. VanRaden Main effects and interactions Transmitting ability Nonadditive merit
Mean value of progeny of random mates Falconer Parents combined value minus parent average Inbreeding depression Best Transmitting Ability Value of progeny of least related mates Value of genes in later generations ADSA 1999 (3) P. M. VanRaden Past and future inbreeding Past inbreeding adjusted yield = yield - (inbreeding)inbreeding)
Future inbreeding adjusted PTA = PTA + (inbreeding)expected inbreeding) combined merit = PTAS + PTAD + (inbreeding)inbreedingS,D) PTAS and PTAD are adjusted for past inbreeding Mating programs ADSA 1999 (4) P. M. VanRaden Test of adjustments Mate population
Data file All Holsteins evaluated August 1998 Regressions 600 females born in 1996 Previously estimated by Wiggans et al. (inbreeding)1995) Timing 4 hours for expected inbreeding; 0 for evaluation
ADSA 1999 (5) P. M. VanRaden PTA Correlations Adjustments None Past Past, Future None 1.0000 .9974 .9976 Past .9974 1.0000 .9976 Past, Future .9976
.9976 1.0000 From 584 active AI bulls ADSA 1999 (6) P. M. VanRaden Value of adjustment Current progress = $200 million/yr Increased progress = 100/(inbreeding)1-.9976) = .24% Yearly benefit = $.5 million/yr Net present value = $10 million/yr
ADSA 1999 (7) P. M. VanRaden Genetic Trend Cows born 1980-1990 kg/year Published trend 104.4 Additive trend 107.4 Additive - inbreeding loss 98.2 ADSA 1999 (8) P. M. VanRaden Interbull Ideal
Proposed Each country adjusts for past inbreeding Do across-country evaluation (inbreeding)MACE) Add future inbreeding within country USA adjusts for past and future inbreeding MACE accounts for change in trend Alternative Adjust only the Net Merit index ADSA 1999 (9)
P. M. VanRaden Conclusions Additive and nonadditive effects can be separated PTAs may include expected nonadditive (inbreeding)expected inbreeding 0) Trends change more than rank Uniform policy best for Interbull ADSA 1999 (10) P. M. VanRaden
Implementation Adjustments easy for U.S. bulls. Foreign pedigree may be missing. Cant add future inbreeding if foreign PTA includes past inbreeding Adjust for past and future in September Interbull test run. Hope MACE converts to new scale. ADSA 1999 (11) P. M. VanRaden
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